Artola A, Bröcher S, Singer W (1990) Different voltage-dependent thresholds for inducing long-term depression and long-term potentiation in slices of rat visual cortex. Nature 347:69-72 [Journal] [PubMed]

Barlow H (2001) Redundancy reduction revisited. Network 12:241-53 [PubMed]

Bell A, Sejnowski T (1995) A non-linear information maximization algorithm that performs blind separation Advances in neural information processing systems, Tesauro G:Touretzky D:Leen T, ed.

Bienenstock EL, Cooper LN, Munro PW (1982) Theory for the development of neuron selectivity: orientation specificity and binocular interaction in visual cortex. J Neurosci 2:32-48 [PubMed]

Bishop C (1995) Neural Networks For Pattern Recognition

Bliss TV, Collingridge GL (1993) A synaptic model of memory: long-term potentiation in the hippocampus. Nature 361:31-9 [Journal] [PubMed]

Dudek SM, Bear MF (1992) Homosynaptic long-term depression in area CA1 of hippocampus and effects of N-methyl-D-aspartate receptor blockade. Proc Natl Acad Sci U S A 89:4363-7 [PubMed]

Fairhall AL, Lewen GD, Bialek W, de Ruyter Van Steveninck RR (2001) Efficiency and ambiguity in an adaptive neural code. Nature 412:787-92 [Journal] [PubMed]

Frey B (1998) Graphical Models for Machine Learning and Digital Communication

Froemke RC, Dan Y (2002) Spike-timing-dependent synaptic modification induced by natural spike trains. Nature 416:433-8 [Journal] [PubMed]

Froemke RC, Tsay IA, Raad M, Long JD, Dan Y (2006) Contribution of individual spikes in burst-induced long-term synaptic modification. J Neurophysiol 95:1620-9 [Journal] [PubMed]

Hebb D (1984) Self-organization and associative memory

Hebb DO (1949) The Organization Of Behavior

Jordan M (1974) Learning in graphical models

Lee TS, Mumford D (2003) Hierarchical Bayesian inference in the visual cortex. J Opt Soc Am A Opt Image Sci Vis 20:1434-48 [PubMed]

Lisman J, Spruston N (1986) From basic neural network principles to neural architecture: Emergence of orientation columns Nature Neurosci 83:839-841

Markram H, Lübke J, Frotscher M, Sakmann B (1997) Regulation of synaptic efficacy by coincidence of postsynaptic APs and EPSPs. Science 275:213-5 [PubMed]

Markram H, Tsodyks M (1996) Redistribution of synaptic efficacy between neocortical pyramidal neurons. Nature 382:807-10 [Journal] [PubMed]

Mongillo M, Deneve S (2006) Online line expectation-maximization in hidden Markov models Unpublished manuscript submitted for publication

Poggio T (1990) A theory of how the brain might work. Cold Spring Harb Symp Quant Biol 55:899-910 [PubMed]

Rabiner L (1989) A tutorial on hidden Markov models and selected applications in speech recognition. Proc IEEE 77:257-286

Senn W, Buchs NJ (2003) Spike-based synaptic plasticity and the emergence of direction selective simple cells: mathematical analysis. J Comput Neurosci 14:119-38 [PubMed]

Sjostrom P, Turrigiano G, Nelson S (1996) Rate, timing, and cooperativity jointly determine cortical synaptic plasticity Neural Comput 8:511-530

Viterbi AJ (1967) Error bounds for convolutional codes and an asymptotically optimum decoding algorithm IEEE Trans Info Theor 13:260-269

Wainwright MJ (1999) Visual adaptation as optimal information transmission. Vision Res 39:3960-74 [PubMed]

Wang HX, Gerkin RC, Nauen DW, Bi GQ (2005) Coactivation and timing-dependent integration of synaptic potentiation and depression. Nat Neurosci 8:187-93 [Journal] [PubMed]

Weiss Y, Freeman WT (2001) Correctness of belief propagation in Gaussian graphical models of arbitrary topology. Neural Comput 13:2173-200 [Journal] [PubMed]

Yao H, Dan Y (2005) Synaptic learning rules, cortical circuits, and visual function. Neuroscientist 11:206-16 [Journal] [PubMed]

Hawkins J, Ahmad S (2016) Why Neurons Have Thousands of Synapses, a Theory of Sequence Memory in Neocortex. Front Neural Circuits 10:23 [Journal] [PubMed]

•	Theory of sequence memory in neocortex (Hawkins & Ahmad 2016) [Model]