/*--------------------------------------------------------------------------
Author: Thomas Nowotny
Institute: Institute for Nonlinear Dynamics
University of California San Diego
La Jolla, CA 92093-0402
email to: tnowotny@ucsd.edu
initial version: 2005-08-17
--------------------------------------------------------------------------*/
#ifndef CN_VALNEURON_CC
#define CN_VALNEURON_CC
#include "CN_neuron.cc"
Valneuron::Valneuron(int inlabel, double *the_p= Val_p):
neuron(inlabel, Val_IVARNO, VALNEURON, the_p, Val_PNO)
{
}
Valneuron::Valneuron(int inlabel, vector<int> inpos, double *the_p= Val_p):
neuron(inlabel, Val_IVARNO, VALNEURON, inpos, the_p, Val_PNO)
{
}
// note that this is the dendritic memb. potential!
inline double Valneuron::E(double *x)
{
assert(enabled);
return x[idx+4];
}
// soma
inline double Valneuron::Esoma(double *x)
{
assert(enabled);
return x[idx];
}
void Valneuron::derivative(double *x, double *dx)
{
static double IVV;
Isyn= 0.0;
forall(den, den_it) {
Isyn+= (*den_it)->Isyn(x);
}
IVV= p[8]*(x[idx]-x[idx+4]);
// differential eqn for E, the membrane potential
dx[idx]= -(pw3(x[idx+1])*x[idx+2]*p[0]*(x[idx]-p[1]) +
pw4(x[idx+3])*p[2]*(x[idx]-p[3])+
p[4]*(x[idx]-p[5])+IVV-p[11])/p[6];
// diferential eqn for m, the probability for one Na channel activation
// particle
_a= 0.32*(-52.0-x[idx]) / (exp((-52.0-x[idx])/4.0)-1.0);
_b= 0.28*(x[idx]+25)/(exp((x[idx]+25)/5.0)-1.0);
dx[idx+1]= _a*(1.0-x[idx+1])-_b*x[idx+1];
// differential eqn for h, the probability for the Na channel blocking
// particle to be absent
_a= 0.128*exp((-48-x[idx])/18.0);
_b= 4.0 / (exp((-25-x[idx])/5.0)+1.0);
dx[idx+2]= _a*(1.0-x[idx+2])-_b*x[idx+2];
// differential eqn for n, the probability for one K channel activation
// particle
_a= .032*(-50-x[idx]) / (exp((-50.0-x[idx])/5.0)-1.0);
_b= 0.5*exp((-55-x[idx])/40.0);
dx[idx+3]= _a*(1.0-x[idx+3])-_b*x[idx+3];
dx[idx+4]= (Isyn + IVV - p[10]*(x[idx+4]-p[5]))/p[9];
}
void Valneuron::noise(double *x, double *dx)
{
dx[idx]= RG.n()*p[10];
for (int i= 1; i < iVarNo; i++) {
dx[idx+i]= 0.0;
}
}
// overloading this one to work on Esoma (!)
void Valneuron::spike_detect(double *x)
{
assert(enabled);
if (Esoma(x) >= SPK_V_THRESH)
{
if (!spiking)
{
start_spiking= 1;
spiking= 1;
spike_time= x[0];
}
else start_spiking= 0;
}
else {
spiking= 0;
start_spiking= 0;
}
}
#endif
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