// genesis
// Creates for each compartment element of L5P an associated randomspike element
// located at the same position, and connected to the L5P cell using volumeconnect.
// The randomspike elements are collected in /Excitatory_fibres.
// The connections may be laid horizontally, using only the z-coordinate as a criterion,
// or radially, allowing the fibre to connect to all compartments at about the same
// position from the soma.
// Inaddition (from 6Oct07) creates for each randomspike element a diffamp from which
// it will receive a RATE msg.
// Moreover, both randomspike and diffamp populations are divided into 3 equal parts:
// feedforward afferents, intracolumn feedbak afferents and intercolumn feedback afferents.
int index
include Fibres.g
make_fibres /Excitatory_fibres
// Connecting fibres to L5P *****
ce /Excitatory_fibres
// First connection scheme: each fiber runs horizontally and is allowed to make a synapse
// on each compartment in (approximately) the same horizontal plane
echo Connecting excitatory feedforward fibres
volumeconnect FF/FF_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-relative \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 1 1 2e-6 \
-probability 0.05 // 1.0 // {P_parallel_fiber_to_Golgi_cell_synapse}
volumeweight FF/FF_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FF/FF_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
echo Connecting excitatory intracolumnar feedback fibres
volumeconnect FBintra/FBintra_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-relative \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 1 1 2e-6 \
-probability 0.05 // 1.0 // {P_parallel_fiber_to_Golgi_cell_synapse}
volumeweight FBintra/FBintra_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FBintra/FBintra_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-fixed 0.02 // -radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
/*
echo Connecting excitatory intercolumnar feedback fibres
volumeconnect FBinter/FBinter_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-relative \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 1 1 2e-6 \
-probability 0.05 // 1.0 // {P_parallel_fiber_to_Golgi_cell_synapse}
volumeweight FBinter/FBinter_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FBinter/FBinter_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
*/
/*
// Second connection scheme: each fiber is allowed to make a synapse on compartments
// located from the soma (the origin) at the same distance as the compartment to which the fibre
// has been associated (remember that a fibre for each compartment was created).
int i
float r, x1, y1, z1
ce {FF}
for (i = 0; i < 686; i = {i} + 1)
x1 = {getfield FF_fibre[{i}] x}
y1 = {getfield FF_fibre[{i}] y}
z1 = {getfield FF_fibre[{i}] z}
r = {sqrt {x1*x1 + y1*y1 + z1*z1}}
// echo {r}
volumeconnect FF_fibre[{i}] \
/L5P/p#[][TYPE=compartment]/AMPA \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 {r + 1e-6} {r + 1e-6} {r + 1e-6} \
-desthole ellipsoid 0 0 0 {r - 1e-6} {r - 1e-6} {r - 1e-6} \
-probability 0.1 // 0.05
end
volumeweight FF_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FF_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
ce {FBintra}
for (i = 0; i < 686; i = {i} + 1)
x1 = {getfield FBintra_fibre[{i}] x}
y1 = {getfield FBintra_fibre[{i}] y}
z1 = {getfield FBintra_fibre[{i}] z}
r = {sqrt {x1*x1 + y1*y1 + z1*z1}}
// echo {r}
volumeconnect FBintra_fibre[{i}] \
/L5P/p#[][TYPE=compartment]/AMPA \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 {r + 1e-6} {r + 1e-6} {r + 1e-6} \
-desthole ellipsoid 0 0 0 {r - 1e-6} {r - 1e-6} {r - 1e-6} \
-probability 0.1 // 0.05
end
volumeweight FBintra_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FBintra_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
ce {FBinter}
for (i = 0; i < 686; i = {i} + 1)
x1 = {getfield FBinter_fibre[{i}] x}
y1 = {getfield FBinter_fibre[{i}] y}
z1 = {getfield FBinter_fibre[{i}] z}
r = {sqrt {x1*x1 + y1*y1 + z1*z1}}
// echo {r}
volumeconnect FBinter_fibre[{i}] \
/L5P/p#[][TYPE=compartment]/AMPA \
-sourcemask box -1 -1 -1 1 1 1 \
-destmask ellipsoid 0 0 0 {r + 1e-6} {r + 1e-6} {r + 1e-6} \
-desthole ellipsoid 0 0 0 {r - 1e-6} {r - 1e-6} {r - 1e-6} \
-probability 0.1 // 0.05
end
volumeweight FBinter_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA -fixed \
{weight_AMPA_synapse} -uniform {weight_distribution}
volumedelay FBinter_fibre[] \
/L5P/p#[][TYPE=compartment]/AMPA \
-radial {E_fibre_conduction_velocity} -uniform {delay_distribution}
*/
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