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Root MJ, MacKinnon R (1994) Two identical noninteracting sites in an ion channel revealed by proton transfer. Science 265:1852-6

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Sack JT, Aldrich RW (2006) Binding of a Gating Modifier Toxin Induces Intersubunit Cooperativity Early in the Shaker K Channel's Activation Pathway J. Gen. Physiol. 128(1):119-132

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