Membrane potential changes in dendritic spines during APs and synaptic input (Palmer & Stuart 2009)

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Accession:120798
" ... Finally, we used simulations of our experimental observations in morphologically realistic models to estimate spine neck resistance. These simulations indicated that spine neck resistance ranges up to ~500 M Ohm. Spine neck resistances of this magnitude reduce somatic EPSPs by ~15%, indicating that the spine neck is unlikely to act as a physical device to significantly modify synaptic strength."
Reference:
1 . Palmer LM, Stuart GJ (2009) Membrane potential changes in dendritic spines during action potentials and synaptic input. J Neurosci 29:6897-903 [PubMed]
Model Information (Click on a link to find other models with that property)
Model Type: Neuron or other electrically excitable cell;
Brain Region(s)/Organism:
Cell Type(s):
Channel(s):
Gap Junctions:
Receptor(s):
Gene(s):
Transmitter(s):
Simulation Environment: NEURON;
Model Concept(s): Active Dendrites; Influence of Dendritic Geometry; Synaptic Integration;
Implementer(s): Morse, Tom [Tom.Morse at Yale.edu]; Stuart, Greg J. [greg.stuart at anu.edu.au];
/*----------------------------------------------------------------
%W%  %G%
ri18um.dat  translated Tue Jun 27 20:56:15 1995 by ntscable 2.0
source file syntax: Nevin
output file syntax: CABLE
soma: diameter = 1 um  length = 0.1 um  area = 0.314159 um2
      3 three-D points numbered 1-3
1 primary neurites
72 branches totaling 8066.41 um in length, 29667.9 um2 in area
1092 tree points translated to 1531 segments (1 requested)
Neurites divided into segments of equal dx between adjacent digitized
points.
Segment length constrained to be < 10 um, or < 10 times
mean diameter of translation interval, whichever is less.
 /*
ri18um.dat, Cell Ri18
coordinates and diameters in Nevin format
pt     type     x       y       z       diam
--------------
Greg's notes

- This cell has modified axon morphology, with the "axon", "soma" and dend1[0] to dend1[7] removed. The bit of axon left is 40 um long and corresponds to the axon initial segment and includes dend1[0] to dend1[4]
- the real soma is dend1[6] to dend1[21]
- dend1[13] is the middle of the soma
- dend1[175] (beginning, ie, 0) is the dendritic recording location
- I added a myelinated axon to the "runRi18_axon.hoc" file
----------------------------------------------------------------*/
                                        /* make geometry explicit */
primary_branches_cell = 1
branches_cell = 72
max_dx_cell = 10
points_cell = 1092
nseg_cell = 1531
double sections_cell[1]
sections_cell[0] = 1091
nsec_cell = 1093
strdef name_section
name_section = "dend"
create    dend1[1083]

/*----------------------------------------------------------------*/
proc geometry() { local i, j
                                        /* neurite geometry*/
    for i = 0,1082 {
        dend1[i] {
		  nseg = fscan()
            pt3dclear()
            for j = 1, fscan() {
                pt3dadd(fscan(),fscan(),fscan(),fscan())
            }
        }
    }

                                        /* branching topology*/
    for i = 1,1082 {
        dend1[fscan()] connect dend1[i] (0), fscan()
    }
}
geometry()

NEURITE COORDINATES AND DIAMETERS:
    1 2
    -0.2    -45.1     -0.6      1.3
    -0.7    -37.3      1.4      1.3
    1 2
    -0.7    -37.3      1.4      1.3
     0.1    -33.9      2.6      1.5
    1 2
     0.1    -33.9      2.6      1.5
     4.1    -27.1      2.4      1.7
    1 2
     4.1    -27.1      2.4      1.7
     4.7    -18.1      2.4        2
    2 2
     4.7    -18.1      2.4        2
       1     -7.8      3.6      2.4
    1 2
       1     -7.8      3.6      2.4
     0.2     -7.5      0.7      4.2
    1 2
     0.2     -7.5      0.7      4.2
     0.2     -5.6      3.4      7.9
    1 2
     0.2     -5.6      3.4      7.9
     0.2     -3.6      3.4       12
    1 2
     0.2     -3.6      3.4       12
     0.2     -2.2      2.3     13.8
    1 2
     0.2     -2.2      2.3     13.8
     0.2        0      4.1     18.9
    1 2
     0.2        0      4.1     18.9
     0.2      0.9      2.5     20.5
    1 2
     0.2      0.9      2.5     20.5
     0.2      3.8      0.4     24.9
    1 2
     0.2      3.8      0.4     24.9
     0.2        5      0.4     26.3
    1 2
     0.2        5      0.4     26.3
     0.2        6      0.4     24.8
    1 2
     0.2        6      0.4     24.8
     0.2      9.7        1     21.9
    1 2
     0.2      9.7        1     21.9
     0.2     14.4      2.1     17.1
    1 2
     0.2     14.4      2.1     17.1
     0.2     16.6      2.1     14.7
    1 2
     0.2     16.6      2.1     14.7
     0.2     19.5      2.1     11.3
    1 2
     0.2     19.5      2.1     11.3
     0.2     20.9      2.1      9.2
    1 2
     0.2     20.9      2.1      9.2
     0.2     22.9      2.1      7.6
    1 2
     0.2     22.9      2.1      7.6
     0.2       29      2.1        7
    1 2
     0.2       29      2.1        7
    -0.5     34.7      3.4      5.6
    1 2
    -0.5     34.7      3.4      5.6
    -1.9       39        4      5.6
    1 2
    -1.9       39        4      5.6
    -0.4     45.4      4.8        5
    1 2
    -0.4     45.4      4.8        5
      -2     49.9      5.1      5.7
    1 2
      -2     49.9      5.1      5.7
     0.6     51.2      4.4      1.2
    1 2
     0.6     51.2      4.4      1.2
     3.6     54.5      4.9        1
    1 2
     3.6     54.5      4.9        1
     3.6     56.8      8.9        1
    1 2
     3.6     56.8      8.9        1
     0.7     57.8      8.6      0.8
    1 2
     3.6     56.8      8.9        1
     6.2     57.1      8.9        1
    1 2
     6.2     57.1      8.9        1
     6.2     59.3      7.5        1
    1 2
     6.2     59.3      7.5        1
     8.7     60.1      8.8      0.6
    1 2
     8.7     60.1      8.8      0.6
    11.4     58.2     11.2      0.6
    1 2
    11.4     58.2     11.2      0.6
    14.5     61.7     11.2      0.6
    1 2
    14.5     61.7     11.2      0.6
    21.1     62.8       10      0.7
    1 2
      -2     49.9      5.1      5.7
    -4.5       59      5.2      5.1
    1 2
    -4.5       59      5.2      5.1
    -4.5     66.6      5.2      5.1
    1 2
    -4.5     66.6      5.2      5.1
    -4.5     72.9      5.2      4.3
    1 2
    -4.5     72.9      5.2      4.3
    -4.9     78.2      5.2      3.7
    1 2
    -4.9     78.2      5.2      3.7
    -4.9       83      4.6        4
    1 2
    -4.9       83      4.6        4
    -8.9     84.9      0.3      1.4
    1 2
    -8.9     84.9      0.3      1.4
   -15.5     84.5     -1.3      1.3
    1 2
   -15.5     84.5     -1.3      1.3
   -19.2     88.6     -2.9      1.2
    1 2
   -19.2     88.6     -2.9      1.2
   -24.3     90.2     -2.9      1.4
    1 2
   -24.3     90.2     -2.9      1.4
   -27.6     94.7     -9.1      1.5
    1 2
   -27.6     94.7     -9.1      1.5
   -34.5     94.7     -6.1      1.6
    2 2
   -34.5     94.7     -6.1      1.6
   -34.5    104.5     -3.8      2.2
    1 2
   -34.5    104.5     -3.8      2.2
   -35.7    109.7     -3.8      1.8
    2 2
   -35.7    109.7     -3.8      1.8
   -43.5    122.2     -9.2      1.4
    1 2
   -43.5    122.2     -9.2      1.4
   -48.3    126.7       -5      1.5
    1 2
   -48.3    126.7       -5      1.5
   -51.6    133.2     -2.9      1.5
    1 2
   -51.6    133.2     -2.9      1.5
   -54.8    138.3     -2.9      1.4
    1 2
   -54.8    138.3     -2.9      1.4
   -56.2    142.6     -4.8      1.9
    2 2
   -56.2    142.6     -4.8      1.9
   -64.3      151     -4.8      1.7
    1 2
   -64.3      151     -4.8      1.7
   -66.4    155.3     -4.8      1.7
    1 2
   -66.4    155.3     -4.8      1.7
   -66.4    161.8     -5.6      1.3
    1 2
   -66.4    161.8     -5.6      1.3
   -68.9    165.5     -4.3        1
    2 2
   -68.9    165.5     -4.3        1
   -79.6    167.6        0      0.9
    1 2
   -79.6    167.6        0      0.9
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    2 2
   -81.5      172        0      0.9
   -87.6    183.6     -3.2      0.8
    1 2
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   -89.9    188.1     -4.6      0.6
    1 2
   -89.9    188.1     -4.6      0.6
   -96.3    193.3     -3.8      0.5
    2 2
   -96.3    193.3     -3.8      0.5
   -96.3    205.1     -1.9      0.7
    2 2
    -4.9       83      4.6        4
    -4.9       95      4.6      3.5
    2 2
    -4.9       95      4.6      3.5
    -4.9    110.1      2.5      3.5
    1 2
    -4.9    110.1      2.5      3.5
    -4.9    112.1      2.5      3.5
    2 2
    -4.9    112.1      2.5      3.5
   -14.5    109.3      3.2      1.6
    1 2
   -14.5    109.3      3.2      1.6
   -20.9    111.1      2.7      1.3
    2 2
   -20.9    111.1      2.7      1.3
   -34.4    106.1      5.9      1.1
    1 2
   -34.4    106.1      5.9      1.1
   -41.1    103.4      6.8      0.9
    1 2
   -41.1    103.4      6.8      0.9
   -43.8    103.7      8.3      0.9
    1 2
   -43.8    103.7      8.3      0.9
   -48.9    102.9      8.3      0.8
    1 2
   -48.9    102.9      8.3      0.8
   -53.9    100.8      5.3      1.1
    1 2
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   -62.1     99.9      6.9      0.8
    1 2
   -62.1     99.9      6.9      0.8
   -66.2     96.6     11.5      0.6
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    1 2
   -67.1     93.6     11.5      0.5
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    1 2
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    1 2
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    -0.3    122.8      0.6        1
    2 2
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     8.5    120.9      5.6      1.2
    1 2
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    18.2    122.6      5.1      0.8
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    1 2
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    1 2
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    1 2
    30.8    135.4      6.8      0.8
    30.8    138.1      7.4        1
    1 2
    30.8    138.1      7.4        1
    38.7    139.4      7.7      1.1
    1 2
    38.7    139.4      7.7      1.1
      48    142.2      6.5      1.3
    2 2
      48    142.2      6.5      1.3
    56.8    142.4     12.3      0.9
    1 2
    56.8    142.4     12.3      0.9
    60.1      145      7.5      0.9
    1 2
    60.1      145      7.5      0.9
    62.2    145.6     12.1      0.4
    1 2
    62.2    145.6     12.1      0.4
    63.3    144.7     13.6      0.4
    1 2
    63.3    144.7     13.6      0.4
    69.1    143.2     13.3      0.4
    1 2
    69.1    143.2     13.3      0.4
    70.1    147.4     13.4      0.7
    1 2
    70.1    147.4     13.4      0.7
    77.4    142.7     17.4      0.6
    1 2
    77.4    142.7     17.4      0.6
      79    142.7     17.4      0.6
    1 2
      48    142.2      6.5      1.3
    53.7    138.3      9.6      0.7
    1 2
    53.7    138.3      9.6      0.7
    59.3    139.9     10.9        1
    2 2
    59.3    139.9     10.9        1
      69    132.5     11.4      0.9
    2 2
      69    132.5     11.4      0.9
    74.8    125.4      8.4      0.6
    2 2
    74.8    125.4      8.4      0.6
    81.6    118.4      5.6      0.5
    1 2
    -2.2    119.3      2.5      3.7
    -2.2    125.6        2      3.9
    1 2
    -2.2    125.6        2      3.9
    -0.3    130.4        2      3.7
    3 2
    -0.3    130.4        2      3.7
    -5.8    152.4      2.9      3.3
    2 2
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    -1.5    162.9      4.9      3.6
    1 2
    -1.5    162.9      4.9      3.6
     1.5      169      4.2      4.2
    1 2
     1.5      169      4.2      4.2
     1.5      169      4.1      1.2
    1 2
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    -3.6    172.2      4.2      0.8
    2 2
    -3.6    172.2      4.2      0.8
   -11.1    179.1     -1.3      1.1
    1 2
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   -13.6      177     -1.3      1.1
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    3 2
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    2 2
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    1 2
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     1.7    -64.2    -16.1      1.1
    1 2
     1.7    -64.2    -16.1      1.1
     1.7    -67.3    -16.1      1.1
    1 2
     1.7    -67.3    -16.1      1.1
     1.7    -70.1    -18.3      1.3
    1 2
     1.7    -70.1    -18.3      1.3
     3.4    -76.9    -18.3      1.3
    1 2
     3.4    -76.9    -18.3      1.3
     3.4    -83.8    -16.5        1
    2 2
     3.4    -83.8    -16.5        1
     7.1    -94.1    -16.5        1
    1 2
     7.1    -94.1    -16.5        1
     5.2    -97.9    -16.5      0.9
    2 2
     5.2    -97.9    -16.5      0.9
     9.8   -107.4    -16.5      0.9
    1 2
     9.8   -107.4    -16.5      0.9
     6.5   -116.8    -16.5        1
    2 2
     6.5   -116.8    -16.5        1
     9.7     -126    -22.4      0.9
    2 2
     9.7     -126    -22.4      0.9
    11.8   -135.8    -24.7      0.8
    2 2
     0.2      0.9      2.5     20.5
    10.4       -4      1.9      1.4
    1 2
    10.4       -4      1.9      1.4
    13.7       -6      1.9      1.2
    1 2
    13.7       -6      1.9      1.2
    20.5     -4.7      1.9      1.5
    1 2
    20.5     -4.7      1.9      1.5
    24.4     -6.6      0.4      1.2
    1 2
    24.4     -6.6      0.4      1.2
    28.6    -13.6      0.4      1.2
    1 2
    28.6    -13.6      0.4      1.2
    35.3    -15.3      0.4      1.5
    2 2
    35.3    -15.3      0.4      1.5
    47.4    -20.2        4      1.5
    1 2
    47.4    -20.2        4      1.5
      51    -19.2        4      1.1
    1 2
      51    -19.2        4      1.1
    53.8    -20.4        4        1
    1 2
    53.8    -20.4        4        1
    57.9    -18.2        6      1.1
    1 2
    57.9    -18.2        6      1.1
    62.9    -18.2        6        1
    1 2
    62.9    -18.2        6        1
    66.2    -17.1        6        1
    1 2
    66.2    -17.1        6        1
    68.5      -18        6      0.7
    1 2
    68.5      -18        6      0.7
    71.2    -17.2        6      0.7
    1 2
    71.2    -17.2        6      0.7
    74.4    -16.6      4.3      0.7
    1 2
    74.4    -16.6      4.3      0.7
    74.4    -14.9      4.3      0.7
    1 2
    74.4    -14.9      4.3      0.7
    76.8    -14.9      4.3      0.7
    1 2
    76.8    -14.9      4.3      0.7
    79.5    -12.5      5.3      0.7
    2 2
    79.5    -12.5      5.3      0.7
    88.6     -7.9      9.4      0.6
    1 2
    88.6     -7.9      9.4      0.6
    92.5     -9.9     12.6      0.6
    2 2
    92.5     -9.9     12.6      0.6
    95.2      -10     -2.2      0.6
    1 2
    47.4    -20.2        4      1.5
    49.6      -23        4      1.2
    1 2
    49.6      -23        4      1.2
    50.7    -30.3        4      1.3
    1 2
    50.7    -30.3        4      1.3
    54.7    -33.1      1.8      0.8
    1 2
    54.7    -33.1      1.8      0.8
      57    -42.7      1.8      1.1
    1 2
      57    -42.7      1.8      1.1
    60.2    -47.4        1        1
    1 2
    60.2    -47.4        1        1
    62.5    -53.7        1        1
    1 2
    62.5    -53.7        1        1
    62.5    -57.6        1        1
    1 2
    62.5    -57.6        1        1
    61.9    -59.7     -0.7        1
    1 2
    61.9    -59.7     -0.7        1
    60.3    -63.3     -0.6      0.8
    1 2
    60.3    -63.3     -0.6      0.8
    65.4    -65.4     -0.6      0.8
    1 2
    65.4    -65.4     -0.6      0.8
    66.2      -72     -0.6      0.8
    1 2
    66.2      -72     -0.6      0.8
    70.7    -75.8     -0.6      0.8
    1 2
    70.7    -75.8     -0.6      0.8
    76.7    -74.2     -0.6      0.8
    1 2
    76.7    -74.2     -0.6      0.8
    82.1      -80     -3.2      0.8
    1 2
    82.1      -80     -3.2      0.8
      87    -78.9     -9.1      0.8
    2 2
      87    -78.9     -9.1      0.8
    95.2    -82.9    -13.7      0.8
    1 2
    95.2    -82.9    -13.7      0.8
    96.8    -86.5    -14.8      0.8
    1 2
    96.8    -86.5    -14.8      0.8
   101.5    -89.4    -14.9      0.7
    3 2
    96.8    -86.5    -14.8      0.8
    90.3    -99.9    -17.6      0.7
    2 2
    61.9    -59.7     -0.7        1
    69.7    -64.7     -0.7      0.7
    2 2
    69.7    -64.7     -0.7      0.7
    75.8    -67.1     -6.2      0.7
    1 2
    75.8    -67.1     -6.2      0.7
    78.8    -69.8     -6.5      0.7
    1 2
    78.8    -69.8     -6.5      0.7
    83.7    -71.4      -11        1
    1 2
    83.7    -71.4      -11        1
    83.7    -71.9      -11        1
    2 2
    83.7    -71.9      -11        1
    90.1    -71.4    -20.6        1
    2 2
    90.1    -71.4    -20.6        1
      99    -76.4    -22.6      0.7
    1 2
      99    -76.4    -22.6      0.7
   105.4    -76.4    -22.6      0.7
    1 2
   105.4    -76.4    -22.6      0.7
   108.5    -79.9    -20.1      0.7
    2 2
     0.2     -2.2      2.3     13.8
    -9.6       -8      2.3      1.7
    1 2
    -9.6       -8      2.3      1.7
    -9.6    -15.8      2.3      1.3
    1 2
    -9.6    -15.8      2.3      1.3
   -12.2    -21.8      2.3      1.3
    1 2
   -12.2    -21.8      2.3      1.3
   -16.8    -22.2      2.3      1.5
    2 2
   -16.8    -22.2      2.3      1.5
     -23    -31.8      2.3      1.7
    2 2
     -23    -31.8      2.3      1.7
   -31.4    -41.7       -3      1.3
    1 2
   -31.4    -41.7       -3      1.3
   -35.6    -43.4       -3      1.3
    1 2
   -35.6    -43.4       -3      1.3
   -40.6    -48.6       -3      1.3
    2 2
   -40.6    -48.6       -3      1.3
   -40.6    -59.9     -1.8      1.3
    2 2
   -40.6    -59.9     -1.8      1.3
   -54.1    -67.8      0.7      1.4
    1 2
   -54.1    -67.8      0.7      1.4
   -60.9    -69.4      0.7      1.2
    1 2
   -60.9    -69.4      0.7      1.2
   -68.7    -72.1      1.9      1.2
    1 2
   -68.7    -72.1      1.9      1.2
   -69.5    -74.9      1.9      1.2
    1 2
   -69.5    -74.9      1.9      1.2
   -69.5      -77        1      0.7
    1 2
   -69.5      -77        1      0.7
   -69.8    -86.2      0.5      0.8
    1 2
   -69.8    -86.2      0.5      0.8
   -66.7    -91.4      0.5      0.8
    1 2
   -66.7    -91.4      0.5      0.8
   -69.7    -95.8      0.5        1
    1 2
   -69.7    -95.8      0.5        1
   -73.4    -97.5      3.1        1
    2 2
   -73.4    -97.5      3.1        1
   -75.1   -109.6      3.1        1
    1 2
   -75.1   -109.6      3.1        1
   -71.8   -112.6     -0.6      0.7
    1 2
   -71.8   -112.6     -0.6      0.7
   -77.2   -118.9     -0.6      0.8
    1 2
   -77.2   -118.9     -0.6      0.8
   -80.8   -123.7      5.8      0.8
    1 2
   -80.8   -123.7      5.8      0.8
   -82.2   -126.4      6.3      0.7
    1 2
   -82.2   -126.4      6.3      0.7
   -84.3     -129        7      0.5
    1 2
   -84.3     -129        7      0.5
   -81.8   -131.5        7      0.5
    1 2
   -81.8   -131.5        7      0.5
   -81.8   -139.5     11.4      0.5
    1 2
   -81.8   -139.5     11.4      0.5
   -85.3   -139.5     13.1      0.5
    1 2
   -85.3   -139.5     13.1      0.5
   -85.3   -142.9     12.3      0.5
    1 2
   -85.3   -142.9     12.3      0.5
   -86.5   -145.6     12.3      0.5
    1 2
   -69.5      -77        1      0.7
   -72.2    -84.1        1        1
    1 2
   -72.2    -84.1        1        1
   -70.6    -87.9        1      0.6
    1 2
   -70.6    -87.9        1      0.6
   -67.6      -93        1      0.6
    1 2
   -67.6      -93        1      0.6
   -71.2    -96.5     -1.3      0.8
    2 2
   -71.2    -96.5     -1.3      0.8
   -75.8   -107.8     -1.9      0.8
    2 2
   -75.8   -107.8     -1.9      0.8
   -81.3   -117.2     -2.3      0.8
    1 2
   -81.3   -117.2     -2.3      0.8
   -86.5   -120.9     -2.3      0.8
    1 2
   -86.5   -120.9     -2.3      0.8
   -90.9   -121.4     -2.3      0.8
    2 2
   -90.9   -121.4     -2.3      0.8
   -93.8   -129.1     -2.3      0.8
    2 2
   -93.8   -129.1     -2.3      0.8
  -101.7   -135.2     -2.3      0.8
    1 2
  -101.7   -135.2     -2.3      0.8
  -108.5   -138.2     -2.3      0.9
    1 2
  -108.5   -138.2     -2.3      0.9
  -108.5   -144.3     -2.3      0.8
    1 2
  -108.5   -144.3     -2.3      0.8
  -111.3   -146.2     -2.3      0.8
    2 2
  -111.3   -146.2     -2.3      0.8
  -115.6   -151.9      2.4      0.8
    1 2
  -115.6   -151.9      2.4      0.8
  -114.8   -154.5      3.5      0.5
    2 2
  -114.8   -154.5      3.5      0.5
  -121.1     -158      7.4      0.5
    2 2
  -121.1     -158      7.4      0.5
  -130.1   -162.5     10.6      0.5
    1 2
  -130.1   -162.5     10.6      0.5
  -132.3   -160.8     10.6      0.5
    2 2
  -132.3   -160.8     10.6      0.5
  -134.2   -169.2     10.3      0.4
    1 2
  -134.2   -169.2     10.3      0.4
  -134.8   -169.2     10.3      0.5
    1 2
     0.2     -3.6      3.4       12
     7.1     -5.8      3.4        2
    1 2
     7.1     -5.8      3.4        2
      11     -4.5      3.4        2
    2 2
      11     -4.5      3.4        2
      11    -12.2     10.3        2
    1 2
      11    -12.2     10.3        2
    16.9    -16.5     10.3        2
    2 2
     0.2     -5.6      3.4      7.9
    -0.5    -17.9      3.4      1.7
    1 2
    -0.5    -17.9      3.4      1.7
    -5.7    -19.8      3.4      1.7
    2 2
    -5.7    -19.8      3.4      1.7
   -16.3    -28.3      4.4      1.6
    1 2
   -16.3    -28.3      4.4      1.6
   -16.7    -29.8       10        1
    1 2
   -16.7    -29.8       10        1
   -13.8    -29.8       10      0.8
    1 2
   -13.8    -29.8       10      0.8
   -14.4    -32.6       10      0.8
    1 2
   -14.4    -32.6       10      0.8
   -22.4    -33.2       10      0.8
    1 2
   -22.4    -33.2       10      0.8
   -30.7    -38.2     12.4      0.5
    1 2
     0.2     -7.5      0.7      4.2
     1.6    -10.1      8.4        2
    1 2
     1.6    -10.1      8.4        2
    -6.4    -10.1      8.4      1.2
    1 2
    -6.4    -10.1      8.4      1.2
   -11.5    -16.4      8.4      0.6
    1 2
   -11.5    -16.4      8.4      0.6
   -14.4    -19.1      9.5      0.6
    2 2
   -14.4    -19.1      9.5      0.6
   -24.9      -14      9.5      0.6
    1 2
   -24.9      -14      9.5      0.6
   -31.2    -18.5      9.5      0.8
    2 2
   -31.2    -18.5      9.5      0.8
   -40.5    -24.7      9.5      0.8
    1 2
   -40.5    -24.7      9.5      0.8
   -43.7      -31      9.5      0.8
    1 2
   -43.7      -31      9.5      0.8
   -46.2    -31.2      9.5      0.8
    1 2
   -46.2    -31.2      9.5      0.8
   -48.2    -28.5      4.6      0.8
    2 2
   -48.2    -28.5      4.6      0.8
   -62.8    -33.7      4.6        1
    1 2
   -62.8    -33.7      4.6        1
     -66    -39.5      4.6      1.1
    1 2
     -66    -39.5      4.6      1.1
   -70.6    -42.6      4.6      1.1
    2 2
   -70.6    -42.6      4.6      1.1
   -81.4    -45.4      4.6      1.1
    1 2
   -81.4    -45.4      4.6      1.1
   -81.4    -43.5        8      1.1
    1 2
   -81.4    -43.5        8      1.1
   -83.9    -39.5        8      0.5
    1 2
   -83.9    -39.5        8      0.5
   -89.9    -35.3        8      0.5
    1 2
   -81.4    -43.5        8      1.1
   -85.8    -47.5      3.9      0.6
    1 2
   -85.8    -47.5      3.9      0.6
   -91.4    -47.5      8.2      0.7
    2 2
   -91.4    -47.5      8.2      0.7
   -98.5    -51.8      8.2      0.7
    1 2
   -98.5    -51.8      8.2      0.7
  -104.9    -50.2      8.2      0.7
    2 2
  -104.9    -50.2      8.2      0.7
  -113.7      -56      9.3      0.7
    1 2
  -113.7      -56      9.3      0.7
  -118.1    -54.9      9.3      0.7
    2 2
  -113.7      -56      9.3      0.7
  -122.2    -59.1      6.1      0.5

CONNECTIONS:	
	
0	1
1	1
2	1
3	1
4	1
5	1
6	1
7	1
8	1
9	1
10	1
11	1
12	1
13	1
14	1
15	1
16	1
17	1
18	1
19	1
20	1
21	1
22	1
23	1
24	1
25	1
26	1
27	1
27	1
29	1
30	1
31	1
32	1
33	1
24	1
35	1
36	1
37	1
38	1
39	1
40	1
41	1
42	1
43	1
44	1
45	1
46	1
47	1
48	1
49	1
50	1
51	1
52	1
53	1
54	1
55	1
56	1
57	1
58	1
59	1
60	1
61	1
39	1
63	1
64	1
65	1
66	1
67	1
68	1
69	1
70	1
71	1
72	1
73	1
74	1
75	1
65	1
77	1
78	1
79	1
80	1
81	1
82	1
83	1
84	1
85	1
86	1
87	1
88	1
89	1
90	1
91	1
92	1
93	1
86	1
95	1
96	1
97	1
98	1
77	1
100	1
101	1
102	1
103	1
104	1
105	1
106	1
107	1
108	1
109	1
110	1
111	1
112	1
107	1
114	1
115	1
116	1
104	1
118	1
119	1
120	1
121	1
122	1
123	1
124	1
125	1
126	1
127	1
128	1
129	1
130	1
131	1
132	1
133	1
121	1
135	1
136	1
137	1
138	1
138	1
140	1
141	1
142	1
143	1
144	1
145	1
146	1
147	1
148	1
149	1
150	1
119	1
152	1
153	1
154	1
153	1
156	1
157	1
158	1
159	1
160	1
161	1
157	1
163	1
164	1
165	1
166	1
167	1
168	1
169	1
170	1
171	1
172	1
173	1
174	1
175	1
176	1
177	1
178	1
179	1
180	1
181	1
182	1
183	1
184	1
185	1
186	1
187	1
180	1
189	1
190	1
191	1
192	1
193	1
194	1
195	1
196	1
197	1
198	1
199	1
200	1
201	1
202	1
203	1
204	1
205	1
206	1
207	1
208	1
209	1
210	1
211	1
212	1
213	1
214	1
215	1
216	1
217	1
218	1
219	1
220	1
221	1
222	1
219	1
224	1
225	1
226	1
227	1
228	1
229	1
230	1
231	1
232	1
233	1
195	1
235	1
236	1
237	1
238	1
238	1
240	1
241	1
242	1
243	1
244	1
245	1
244	1
247	1
248	1
191	1
250	1
251	1
252	1
253	1
254	1
255	1
256	1
257	1
258	1
259	1
260	1
261	1
262	1
263	1
264	1
265	1
266	1
267	1
268	1
269	1
270	1
271	1
272	1
273	1
274	1
275	1
276	1
277	1
278	1
279	1
280	1
281	1
282	1
283	1
284	1
285	1
286	1
287	1
288	1
289	1
290	1
291	1
292	1
293	1
293	1
295	1
296	1
297	1
298	1
299	1
300	1
301	1
289	1
303	1
304	1
305	1
306	1
279	1
308	1
309	1
310	1
311	1
312	1
313	1
314	1
315	1
316	1
317	1
317	1
319	1
320	1
321	1
321	1
323	1
258	1
325	1
326	1
327	1
328	1
329	1
330	1
331	1
332	1
333	1
334	1
335	1
336	1
337	1
338	1
339	1
340	1
341	1
342	1
343	1
344	1
345	1
346	1
347	1
348	1
349	1
350	1
351	1
352	1
353	1
352	1
355	1
356	1
357	1
358	1
359	1
330	1
361	1
362	1
363	1
364	1
365	1
366	1
367	1
368	1
369	1
370	1
371	1
372	1
373	1
374	1
375	1
376	1
377	1
378	1
379	1
380	1
381	1
382	1
383	1
383	1
378	1
386	1
387	1
388	1
389	1
390	1
391	1
392	1
393	1
394	1
395	1
396	1
176	1
398	1
399	1
400	1
401	1
402	1
403	1
404	1
405	1
406	1
407	1
408	1
409	1
410	1
411	1
412	1
413	1
414	1
415	1
416	1
417	1
418	1
419	1
420	1
421	1
422	1
423	1
424	1
425	1
426	1
427	1
428	1
429	1
430	1
431	1
432	1
433	1
434	1
435	1
436	1
437	1
438	1
439	1
440	1
441	1
442	1
443	1
440	1
445	1
446	1
447	1
448	1
449	1
450	1
451	1
452	1
430	1
454	1
455	1
456	1
457	1
458	1
459	1
460	1
461	1
462	1
463	1
464	1
465	1
466	1
467	1
468	1
469	1
470	1
417	1
472	1
473	1
474	1
475	1
476	1
477	1
478	1
479	1
480	1
481	1
482	1
483	1
484	1
485	1
486	1
487	1
488	1
489	1
490	1
491	1
492	1
493	1
494	1
495	1
496	1
481	1
498	1
499	1
500	1
501	1
502	1
503	1
504	1
505	1
506	1
507	1
508	1
509	1
510	1
511	1
512	1
513	1
514	1
515	1
516	1
517	1
506	1
519	1
520	1
521	1
522	1
523	1
524	1
525	1
526	1
527	1
528	1
529	1
530	1
531	1
399	1
533	1
534	1
535	1
536	1
537	1
538	1
539	1
540	1
541	1
542	1
543	1
544	1
545	1
546	1
547	1
548	1
549	1
550	1
551	1
552	1
553	1
554	1
555	1
556	1
557	1
558	1
559	1
560	1
561	1
562	1
563	1
564	1
565	1
566	1
567	1
568	1
569	1
570	1
571	1
572	1
573	1
574	1
575	1
576	1
577	1
578	1
579	1
580	1
581	1
582	1
583	1
584	1
585	1
586	1
567	1
588	1
589	1
590	1
591	1
592	1
593	1
594	1
595	1
596	1
597	1
598	1
599	1
600	1
601	1
602	1
603	1
604	1
605	1
606	1
607	1
608	1
609	1
610	1
611	1
612	1
613	1
614	1
615	1
616	1
617	1
618	1
619	1
620	1
621	1
622	1
623	1
624	1
625	1
626	1
553	1
628	1
629	1
630	1
631	1
632	1
633	1
634	1
635	1
636	1
637	1
638	1
639	1
640	1
641	1
642	1
643	1
644	1
645	1
646	1
647	1
648	1
649	1
650	1
651	1
652	1
653	1
654	1
655	1
656	1
657	1
658	1
659	1
660	1
661	1
662	1
663	1
664	1
665	1
666	1
667	1
668	1
669	1
670	1
671	1
672	1
673	1
674	1
675	1
676	1
677	1
633	1
679	1
680	1
681	1
682	1
683	1
684	1
685	1
686	1
682	1
688	1
689	1
690	1
691	1
692	1
693	1
694	1
695	1
696	1
697	1
698	1
699	1
700	1
701	1
702	1
703	1
704	1
705	1
706	1
707	1
708	1
709	1
710	1
711	1
712	1
713	1
714	1
715	1
716	1
717	1
718	1
719	1
720	1
721	1
722	1
723	1
724	1
725	1
726	1
727	1
728	1
729	1
730	1
731	1
722	1
733	1
734	1
735	1
736	1
737	1
738	1
739	1
720	1
741	1
742	1
743	1
744	1
745	1
746	1
747	1
748	1
749	1
750	1
22	1
752	1
753	1
754	1
755	1
756	1
757	1
758	1
759	1
760	1
761	1
762	1
763	1
753	1
765	1
766	1
767	1
13	1
769	1
770	1
771	1
772	1
773	1
774	1
775	1
776	1
777	1
778	1
779	1
780	1
781	1
782	1
783	1
776	1
785	1
786	1
787	1
788	1
789	1
790	1
775	1
792	1
793	1
794	1
795	1
796	1
797	1
798	1
12	1
800	1
801	1
802	1
803	1
804	1
805	1
806	1
807	1
808	1
803	1
810	1
811	1
812	1
813	1
814	1
815	1
816	1
817	1
818	1
819	1
820	1
821	1
822	1
823	1
817	1
825	1
826	1
813	1
828	1
829	1
830	1
831	1
832	1
833	1
834	1
835	1
836	1
837	1
838	1
839	1
840	1
841	1
842	1
843	1
844	1
845	1
846	1
847	1
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849	1
850	1
851	1
852	1
853	1
854	1
855	1
856	1
857	1
810	1
859	1
860	1
861	1
862	1
863	1
864	1
865	1
866	1
867	1
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871	1
872	1
802	1
874	1
875	1
876	1
877	1
878	1
879	1
880	1
881	1
882	1
883	1
884	1
885	1
886	1
887	1
888	1
889	1
890	1
885	1
892	1
893	1
894	1
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896	1
897	1
898	1
892	1
900	1
901	1
902	1
903	1
904	1
905	1
906	1
907	1
908	1
909	1
910	1
911	1
912	1
913	1
11	1
915	1
916	1
917	1
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916	1
920	1
921	1
922	1
920	1
924	1
925	1
926	1
927	1
928	1
929	1
930	1
931	1
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933	1
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935	1
927	1
937	1
938	1
939	1
940	1
941	1
942	1
943	1
944	1
945	1
946	1
947	1
10	1
949	1
950	1
951	1
952	1
953	1
954	1
955	1
956	1
957	1
958	1
959	1
960	1
961	1
962	1
963	1
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965	1
966	1
967	1
968	1
955	1
970	1
971	1
972	1
973	1
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975	1
976	1
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978	1
979	1
980	1
981	1
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985	1
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986	1
977	1
989	1
990	1
991	1
992	1
993	1
994	1
995	1
996	1
8	1
998	1
999	1
1000	1
1001	1
1002	1
1003	1
1004	1
1005	1
1006	1
1007	1
1008	1
1009	1
1010	1
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1015	1
1016	1
1017	1
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1020	1
1021	1
1022	1
1023	1
1024	1
1025	1
1011	1
1027	1
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1030	1
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1034	1
1035	1
1036	1
1037	1
1038	1
1039	1
1040	1
1041	1
1042	1
1043	1
1044	1
1045	1
7	1
1047	1
1048	1
1049	1
6	1
1051	1
1052	1
1053	1
1054	1
1055	1
1056	1
1057	1
5	1
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1060	1
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1063	1
1064	1
1065	1
1066	1
1067	1
1068	1
1069	1
1070	1
1071	1
1072	1
1073	1
1074	1
1073	1
1076	1
1077	1
1078	1
1079	1
1080	1
1080	1
	

proc correct() {
  /* correct diameters of dendrites at origins of basals */
        dend1[769] {diam=1.40}
        dend1[800] {diam=2.40}
        dend1[915] {diam=1.40}
        dend1[949] {diam=1.40}
        dend1[998] {diam=1.70}
        dend1[1047] {diam=2.00}
        dend1[1051] {diam=1.70}
        dend1[1059] {diam=2.00}
  /* correct diameters of dendrites at origins of obliques */
        dend1[25] {diam=1.20}
        dend1[40] {diam=1.35}
        dend1[66] {diam=1.60}   
        dend1[78] {diam=2.45}
        dend1[105] {diam=0.80}
        dend1[120] {diam=1.00}
        dend1[154] {diam=0.80}
        dend1[752] {diam=1.30}
}
correct()
/*----------------------------------------------------------------*/
proc geometry() { 
        /* NULL geometry procedure: keeps the user from
        calling the geometry procedure in isolation. */
    printf("\nYou must re-read the entire geometry\n")
    printf("file to execute geometry().\n\n")
}


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