A two population spiking continuous attractor model of grid cells. This model combines the attractor dynamics with theta-nested gamma oscillatory activity. It reproduces the behavioural response of grid cells (grid fields) in medial entorhinal cortex, while at the same time allowing for nested gamma oscillations of post-synaptic currents.
Pastoll H, Solanka L, van Rossum MC, Nolan MF (2013) Feedback inhibition enables theta-nested gamma oscillations and grid firing fields. Neuron 77:141-54 [PubMed]